Ole Seberg, Laboratory of Molecular Systematics, Natural History Museum of Denmark (Supervisor)

Gitte Petersen, Laboratory of Molecular Systematics, Natural History Museum of Denmark (Supervisor)

H. Peter Linder, Institute of Systematic Botany, Zurich, Switzerland (Co-supervisor)

Nina Rønsted, Faculty of Pharmaceutical Sciences, University of Copenhagen

John and Sandie Burrows, Buffelskloof Private Nature Reserve, South Africa